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During this stage the nose is the primary sense organ used by puppies to find their mother's teats, and to locate their littermates, if they become separated by a short distance.  Puppies develop very quickly during their first three months, particularly after their eyes and ears open and they are no longer completely dependent on their mother.  If the litter exceeds six puppies, particularly if one or more are obvious runts, human intervention in hand-feeding the stronger puppies is necessary to ensure that the runts get proper nourishment and attention from the mother.  Born after an average of 63 days of gestation, puppies emerge in an amnion that is bitten off and eaten by the mother dog.  The mother may regurgitate partially digested food for the puppies or might let them eat some of her solid food.  In addition, puppies' ears remain sealed until about thirteen to seventeen days after birth, after which they respond more actively to sounds.  Puppies open their eyes about nine to eleven days following birth.  Puppies are born with a fully functional sense of smell but can't open their eyes.  As they reach one month of age, puppies are gradually weaned and begin to eat solid food.  Between two and four weeks old, puppies usually begin to growl, bite, wag their tails, and bark.  At first, puppies spend the large majority of their time sleeping and the rest feeding.  Puppies begin to nurse almost immediately.  Puppies are not able to see as well as adult dogs.  They instinctively pile together into a heap, and become distressed if separated from physical contact with their littermates, by even a short distance.  In vernacular English, puppy refers specifically to dogs, while pup may often be used for other mammals such as seals, giraffes, guinea pigs, or even rats or sharks.  The mother dog usually refuses to nurse at this stage, though she might let them occasionally nurse for comfort.  Their coordination and strength improve, they spar with their littermates, and begin to explore the world outside the nest.  A puppy's coat color may change as the puppy grows older, as is commonly seen in breeds such as the Yorkshire Terrier.  During their first two weeks, a puppy's senses all develop rapidly.  At first, their retinas are poorly developed and their vision is poor.  They play wrestling, chase, dominance, and tug-of-war games.